Evolutionary Ecology

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Niels Dingemanse

Prof. Dr. Niels Dingemanse


Phone: +49 89 2180 74 209

Room B02.008

LMU München/Department Biology II
Behavioural Ecology
Großhaderner Str. 2
82152 Planegg-Martinsried

Phone: Tel: +49 (0) 89 2180 74 209
Fax: +49 (0) 89 2180 74 204



For more details on our current research at the LMU see the webpage “Research”.



Personal information:

Birth date: 13 March 1974, Tholen (The Netherlands)

Citizenship: The Netherlands

Marital Status: married, three children


  • 1992: “Atheneum” at Comenius College, Hilversum (The Netherlands)
  • 1997: M.Sc (Honours) in Ecology, University of Groningen (the Netherlands)
  • 2003: PhD in natural sciences, University of Utrecht (the Netherlands)

Academic Positions:

  • 2012-present: Tenured Professor in Behavioural Ecology, Department of Biology , Ludwig Maximilians University, Germany
  • 2012-2017: Head of Research Group (Evolutionary Ecology of Variation), Max Planck Institute for Ornithology, Seewiesen, Germany
  • 2009-2012: Research scientist at the Department of Behavioural Ecology and Evolutionary Genetics, MPIO Seewiesen
  • 2006-2009: Postdoctoral Research Fellow, University of Groningen, The Netherlands (fellowship within the Innovational Research Incentives Scheme Veni of the Netherlands Organisation for Scientific Research)
  • 2005-2006: Postdoctoral Research Fellow, University of Groningen, The Netherlands (ALW open competition grant of the Netherlands Organisation for Scientific Research)
  • 2004: Postdoctoral Research Fellow, University of Groningen, The Netherlands (Honorary scholarship of the University of Groningen)
  • 2003-2004: Postdoctoral Research Fellow, University of Wales, Bangor, UK (TALENT scholarship of the Netherlands Organisation for Scientific Research)
  • 1998-2003: PhD at Netherlands Institute of Ecology, Heteren, the Netherlands (Advisors: Dr. P.J. Drent and Prof. A.J. van Noordwijk) The thesis is freely available as a pdf-file at: http://igitur-archive.library.uu.nl/dissertations/2003-1218-101459/inhoud.htmtop


In Press

Dingemanse N.J., Wright J. (2020). Criteria for acceptable studies of animal personality and behavioural syndromes. Ethology. In Press.

Schielzeth H., Dingemanse N.J., Nakagawa S., Westneat D.F., Allegue H., Teplitsky C., Réale D., Dochtermann N.A., Garamszegi L.Z., Araya-Ajoy Y.G. (2020). Robustness of linear mixed-effects models to violations of distributional assumptions. Methods in Ecology and Evolution. In Press.

Han C., Brooks R., Dingemanse N.J. (2020). Condition-dependent mutual mate preference and intersexual genetic correlations for mating activity. American Naturalist. In Press.

Wetzel D.P., Mutzel A., Wright J., Dingemanse N.J. (2020). Novel sources of (co)variation in nestling begging behavior and hunger. Behav. Ecol. In Press.

In peer-reviewed journals


Dingemanse N.J., Barber I, & Dochtermann N.A. (2020). Non-consumptive effects of predation: does perceived risk strengthen the genetic integration of behaviour and morphology in stickleback? Ecology Letters 23:107-118.

Dingemanse N.J., Moiron M., Araya-Ajoy Y.G., Mouchet A. & Abbey-Lee R.N. (2020). Individual variation in age-dependent reproduction: Fast explorers live fast but senesce young? Journal of Animal Ecology 89:601-613.

Hertel A.G., Niemelä P.T., Dingemanse N.J., Mueller T. (2020). A guide for studying among-individual behavioral variation from movement data in the wild. Movement Ecology 8:30.

Merten von S, Dingemanse N.J., Mathias M.D. & Rychlik L. (2020). Individual behavior, behavioral stability, and pace of life within and among five shrew species. Behavioral Ecology and Sociobiology 74(2):15.

Strauss A.F.T., Hutfluss A., Dingemanse N.J. (2020). Great tits responding to territorial intrusions sing less but alarm more on colder days. Ethology 126: 445-454.



Abbey-Lee R.N., Dingemanse N.J. 2019. Adaptive individual variation in phenological responses to perceived predation levels. Nature Communications, 10:1601.

Mathot K.J., Dingemanse N.J., & Nakagawa S. 2019. The covariance between metabolic rate and behaviour varies across behaviours and thermal types: meta-analytic insights. Biological Reviews, 94: 1056-1074.

Niemela P.T., Niehoff P., Gasparin C., Dingemanse N.J., Tuni C. 2019. Crickets become behaviourally more stable when raised under higher temperatures. Behavioral Ecology and Sociobiology: 73-81.

Wang D., Forstmeier W., Valcu M., Dingemanse N.J., Bulla M., Both C., Duckworth R.A., Kiere L.M., Karell K., Albrecht T., Kempenaers B. 2019. Scrutinizing assortative mating in birds. PLOS Biology, 17: e3000156.

Wright J., Boldstad G., Araya-Ajoy Y.G., & Dingemanse N.J. 2019. Life-history evolution under fluctuating density-dependent selection and the adaptive alignment of pace-of-life-syndromes. Biological Reviews 94, 230-247.



Abbey-Lee R., Araya-Ajoy Y.G., Mouchet A., Moiron M; Stuber E; Kempenaers B & Dingemanse N.J. 2018. Does perceived predation risk affect patterns of extra-pair paternity? A field experiment in a passerine bird. Functional Ecology 32:1001-1010.

Araya-Ajoy Y.G., Boldstad G.H., Brommer J., Careau V., Dingemanse N.J., Wright J. 2018. Demographic measures of an individual's "pace of life": fecundity rate, lifespan, generation time or a composite variable? Behavioral Ecology and Sociobiology 72:UNSP 75.

Dammhahn M., Dingemanse N.J., Niemelä P.T., Réale D. 2018. Pace-of-life syndromes: a framework for the adaptive integration of behaviour, physiology and life history, Behavioral Ecology and Sociobiology, 72:62.

Gasparini C., Lu C., Dingemanse N.J, Tuni C. 2018. Paternal-effects in a terrestrial ectotherm are temperature dependent but no evidence for adaptive effects. Functional Ecology 32(4):1011-1021.

Han CS, Tuni C, Ulcik J, & Dingemanse NJ. 2018. Increased developmental density decreases the magnitude of indirect genetic effects expressed during agonistic interactions in an insect. Evolution 72:2435-2448.

Moiron M., Mathot K.J & Dingemanse N.J. 2018. To eat and not be eaten: diurnal mass gain and foraging strategies in wintering great tits. Proceedings of the Royal Society B-Biological Sciences 1874: 20172868.

Niemela P.T., Dingemanse N.J. 2018. Meta-analysis reveals weak associations between intrinsic state and personality. Proceedings B, 285: 20172823.

Niemelä PT & Dingemanse NJ. 2018. On the usage of single measurements in behavioural ecology research on individual differences. Animal Behaviour 145:99-105.

Tuni C, Han CS, & Dingemanse NJ. 2018. Multiple biological mechanisms result in correlations between pre- and post-mating traits that differ among versus within individuals and genotypes. Proceedings of the Royal Society B-Biological Sciences 285(1885).

van Overveld T, et al. 2018. Food predictability and social status drive individual resource specializations in a territorial vulture. Scientific Reports 8:15155.



Allegue H., Araya-Ajoy Y.G., Dingemanse N.J., Dochtermann N.A., Garamszegi L.Z., Nakagawa S., Réale D., Schielzeth H. & Westneat D.F. 2017. Statistical Quantification of Individiual Differences (SQuID): an educational and statistical tool for understanding multilevel phenotypic data in linear mixed models. Methods in Ecology and Evolution, 8, 257-267.

Araya-Ajoy Y.G. & Dingemanse N.J. 2017. Repeatability, heritability, and age-dependence of seasonal plasticity in aggressiveness in a wild passerine bird. Journal of Animal Ecology, 86, 227-238.

Class B., Dingemanse N.J., Araya-Ajoy Y.G., Brommer J.E. 2017. A statistical methodology for estimating assortative mating for phenotypic traits that are labile or measured with error. Methods in Ecology and Evolution, 8, 1910-1919.

Dingemanse N.J. 2017. The role of personality research in contemporary behavioral ecology: a comment on Beekman and Jordan. Behavioral Ecology, 28, 624-625.

Han C.S. & Dingemanse N.J. 2017. You are what you eat: diet shapes body composition, personality, and behavioural stability. BMC Evolutionary Biology, 17, 8.

Han C.S. & Dingemanse N.J. 2017. Protein deprivation decreases male survival and the intensity of sexual antagonism in Southern field crickets Gryllus bimaculatus. Journal of Evolutionary Biology, 30, 839-847.

Han C.S. & Dingemanse N.J. 2017. Sex-dependent expression of behavioural genetic architectures and the evolution of sexual dimorphism. Proceedings B, 284, 1864.

Kelleher S.R., Silla A.J., Dingemanse N.J. , Byrne P. Body size predicts between-individual differences in exploration behaviour in the southern corroboree frog. Animal Behaviour, 129, 161-170.

Mathot K.J., Olsen A.-L., Mutzel A., Araya-Ajoy Y.G., Nicolaus M., Westneat D, Wright J., Kempenaers B., Dingemanse N.J. 2017. Provisioning tactics of great tits in response to long-term brood manipulations differ across years. Behavioral Ecology, 28, 1402-1413.

Niemelä P.T. & Dingemanse N.J. 2017. Trustworthiness of online beer ratings as a source of social information. Behavioral Ecology and Sociobiology, 61, 24.

Niemelä P.T., Dingemanse N.J. 2017. Individual versus pseudo-repeatability in behaviour: lessons from translocation experiments in a wild insect. Journal of Animal Ecology, 86, 1033-1043.

Niemelä P.T. & Dingemanse N.J. 2017. Trustworthiness of online beer ratings as a source of social information. Behavioral Ecology and Sociobiology, 71, 24.

Santostefano F., Wilson A.J., Niemelä P.T., Dingemanse N.J. 2017. Indirect genetic effects: a key component of the genetic architecture of behaviour. Scientific Reports, 7, 10235.

Santostefano F., Wilson A.J., Niemelä P.T., Dingemanse N.J. 2017. Behavioural mediators of genetic life-history trade-offs: a test of the pace-of-life syndrome hypothesis in field crickets. Proceedings B, 284, 1864.

Senar J.C., Garamszegi L.Z., Tilgar V., Biard C., Moreno-Rueda G., Salmón P., Rivas J.M., Sprau P., Dingemanse N.J., Charmantier A. 2017. Urban great tits (Parus major) show higher distress calling and pecking rates than rural birds across Europe. Frontiers in Ecology and Evolution, 5, 163.

Serrano-Davies E., Araya-Ajoy Y.G., Dingemanse N.J., Sanz J.J. 2017. Personality-related differences in response to habitat in Mediterranean blue tits. Ethology, 123, 861-869.

Sprau P. & Dingemanse N.J. 2017. An approach to distinguish between plasticity and non-random distributions of behavioural types along urban gradients in a wild passerine bird. Frontiers in Ecology and Evolution, 5, 92.

Sprau P., Mouchet A., Dingemanse N.J. 2017. Multi-dimensional environmental predictors of variation in avian forest and city life-histories. Behavioral Ecology, 28, 59-68.

Stuber E., Dingemanse N.J., Mueller J. 2017. Temperature affects frequency but not rhythmicity of nocturnal awakenings in free-living great tits. Animal Behaviour, 135-141.

Vincze E., Seress G., Lagisz M., Nakagawa S., Dingemanse N.J., Sprau P. Does urbanization affect predation of bird nests? A meta-analysis. Frontiers in Ecology and Evolution, 5, 29.



Abbey-Lee R., Kaiser A., Mouchet A., Dingemanse N.J. 2016. Immediate and carry-over effects of perceived predation risk on communication behavior in wild birds. Behavioral Ecology, 27, 708-716.

Abbey-Lee R., Mathot K.M., Dingemanse N.J. 2016. Behavioral and morphological responses to perceived predation risk: a field experiment in passerines. Behavioral Ecology, 27, 857-864.

Araya-Ajoy Y.G., Dingemanse N.J. & Kempenaers, B. 2016. Timing of extra-pair fertilizations: trade-offs or pair synchrony spill-overs? Behavioral Ecology, 27, 377-384 .

Araya-Ajoy Y.G., Kuhn S., Mathot K.J., Mouchet A., Mutzel A., Nicolaus M., Wijmenga J., Kempenaers B., Dingemanse N.J. 2016. Sources of (co)variation in the alternative siring routes available to male great tits (Parus major). Evolution, 70, 2308-2321.

Berger-Tal R., Berner-Aharon N., Aharon S. Tuni C. & Lubin Y. 2016. Good reasons to leave home: proximate dispersal cues in a social spider. Journal of Animal Ecology, 85, 1035-1042.

Han C.S. & Jablonski P.G. 2016. Predators induce conditions for size-dependent alternative reproductive tactics in a water strider male. Animal Behaviour, 111, 271-279.

Han C.S., Jaeger H. & Dingemanse N.J. 2016. Individuality in nutritional preferences: a multi-level approach in field crickets. Scientific Reports, 6, 29071.

Han C.S., Santostefano, F. & Dingemanse, N.J. 2016. Do social partners affect same-sex behaviour in male water striders? Animal Behaviour, 116, 53-59.

Mathot K.J., Abbey-Lee R.N., Kempenaers B., Dingemanse N.J. 2016. Do great tits (Parus major) suppress basal metabolic rate in response to increased perceived predation danger? A field experiment. Physiology & Behaviour, 164, 400-406.

Mils JA. et al. 2016. Solutions for archiving data in long-term studies: A Reply to Whitlock et al. Trends in Ecology and Evolution, 31, 85-87.

Moiron M., Mathot K.J. & Dingemanse N.J. 2016. A multi-level approach to quantify speed-accuracy trade-offs in great tits (Parus major). Behavioral Ecology, 27, 1539-1546.

Nicolaus M., Piault R., Ubels R., Tinbergen J.M. & Dingemanse N.J. 2016. The correlation between colouration and exploration behaviour varies across hierarchical levels in a wild passerine bird. Journal of Evolutionary Biology, 29, 1780-1792.

Nicolaus M., Tinbergen J.M., Ubels R., Both C. & Dingemanse N.J. 2016. Density fluctuations represent a key process maintaining personality variation in a wild passerine bird. Ecology Letters, 19, 478-486.

Santostefano F., Wilson A.J., Araya-Ajoy Y.G. & Dingemanse N.J. 2016. Interacting with the enemy: indirect effects of personality on conspecific aggression in field crickets. Behavioral Ecology, 24, 1235-1246.

Stuber E.F., Baumgarter C., Dingemanse N.J., Kempenaers B. & Mueller J.C. 2016. Genetic correlates of individual differences in sleep behaviour in free-living great tits (Parus major). G3: Genes, Genomes, Genetics, 6, 599-607.

Tuni C., Perdigón Ferreira J., Fritz Y., Munoz Meneses A. & Gasparini C. 2016. Impaired sperm quality and delayed mating but no costs for offspring fitness in field crickets winning a fight. Journal of Evolutionary Biology, 29, 1643-1647.



Araya-Ajoy Y.G., Mathot, K.M., & Dingemanse N.J. 2015. An approach to estimate short-term, long-term and reaction norm repeatability. Methods in Ecology and Evolution.

Dingemanse N.J. & Araya-Ajoy Y.G. 2015. Interacting personalities: behavioural ecology meets quantitative genetics. Trends in Ecology and Evolution, 30, 88-97.

Hall M.L., van Asten T.T., Katsis A.C., Dingemanse N.J. Magrath M.J.L. & Mulder R.A. 2015. Animal personality and pace-of-life syndromes: do fast-exploring fairy-wrens die young? Frontiers in Ecology and Evolution, 3, 28.

Han C.S. & Dingemanse N.J. 2015. Effect of diet on the structure of animal personality. Frontiers in Zoology 12(Suppl 1), S5.

Mathot K.J. & Dingemanse N.J. 2015. Energetics and behaviour: reply to Careau and Garland. Trends in Ecology and Evolution, 30, 367-368.

Mathot K.J. & Dingemanse N.J. 2015. Studies linking energetics and behaviour: unrequited needs and new directions. Trends in Ecology and Evolution, 30, 199-206.

Mathot K.J. & Dingemanse N.J. 2015. Personality and Plasticity. In: Integrative Organismal Biology (Ed. by Martin, L.B., Ghalambor, C.K. & Woods, H.A.) Willey Scientific, pp. 55-69.

Mathot K.J., Nicolaus M., Araya-Ajoy Y.G., Dingemanse N.J. & Kempenaers B. 2015. Does metabolic rate predict risk-taking behaviour: a field experiment in a wild passerine bird. Functional Ecology 29, 239-249.

Niemelä P., Lattenkamp E.Z. & Dingemanse N.J. 2015. Personality-related survival and sampling bias in wild cricket nymphs. Behavioral Ecology, 26, 936-946.

Sih A., Mathot K.J., Moirón M., Montiglio P.O., Wolf M. & Dingemanse N.J. 2015. Animal personality and state-behaviour feedbacks: a review and guide for empiricists. Trends in Ecology & Evolution, 30, 50-60.

Stuber E.F., Mathot K.J., Kempenaers B., Dingemanse N.J., Mueller J.C. 2015. Sex-specific association between sleep and basal metabolic rate in great tits. Animal Behaviour, 109, 15-22.

Stuber E.F., Dingemanse N.J., Kempenaers B., Mueller J.C. 2015. Sources of intraspecific variation in sleep behaviour of wild great tits (Parus major). Animal Behaviour, 106, 201-221.

Westneat D.F., Wright J. & Dingemanse N.J. 2015. The biology hidden inside residual within-individual phenotypic variation. Biological Reviews, 90, 729-743.



Araya-Ajoy J.G. & Dingemanse N.J. 2014. Characterizing behavioural 'characters': an evolutionary framework. Proceedings of the Royal Society of London Series B, 281, 20132645

Baugh A.T., van Oers K., Dingemanse N.J. & Hau E. 2014. Baseline and stress-induced glucocorticoid concentrations are not repeatable but covary within individual great tits (Parus major). General and Comparative Endocrinology, 208, 154-163.

Dingemanse N.J. & Dochtermann N.A. 2014. Individual Behaviour: Behavioural Ecology meets Quantitative Genetics. In: Quantitative Genetics in the Wild (Ed. by Charmantier, A., Garant, D. & Kruuk, L.E.B.) Oxford University Press, pp. 54-67.

Mueller J.C., Edelaar P., Carrete M., Serrano D., Potti J., Blas J., Dingemanse N.J., Kempenaers B. & Tella J.L. 2014. Behaviour-related DRD4 polymorphisms in invasive bird populations. Molecular Ecology, 23, 2876-2885.

Nicolaus M., Mathot K.J., Araya-Ajoy Y.G., Mutzel A., Wijmenga J.J., Kempenaers B. & Dingemanse N.J. 2014. Does coping style predict optimization: an experimental test in a wild passerine bird. Proceedings of the Royal Society of London Series B, 282, 20142405.

Niemelä P.T. & Dingemanse N.J. 2014. Artificial environments and the study of ‘adaptive’ personalities. Trends in Ecology and Evolution, 29, 245-247.

Stuber E.F., Grobis M.M., Abbey-Lee R., Kempenaers B., Mueller J.C. & Dingemanse N.J.2014 Perceived predation risk affects sleep behaviour in free-living great tits, Parus major. Animal Behaviour, 98, 157-165.



Mutzel A., Dingemanse N.J., Araya-Ajoy Y.G. & Kempenaers B. 2013. Parental provisioning behaviour plays a key role in linking personality with reproductive success. Proceedings of the Royal Society B-Biological Sciences, 280, 20131019

Araya-Ajoy J.G. & Dingemanse N.J. 2013. Characterizing behavioural 'characters': an evolutionary framework. Proceedings of the Royal Society of London Series B, 281, 20132645.

Dingemanse N.J. & Dochtermann N.A. 2013. Quantifying individual variation in behaviour: mixed-effect modelling approaches. Journal of Animal Ecology, 82, 39-54.

Dingemanse N.J. & Wolf M. 2013. Between-individual differences in behavioural plasticity: causes and consequences. Animal Behaviour, 85, 1031-1039.

Dingemanse N.J. & Réale D. 2013. What is the evidence for natural selection maintaining animal personality variation? In: Animal Personalities: Behaviour, Physiology, and Evolution (Ed. by Carere, C. & Maestripieri, D.) Chicago: Chicago University Press.

Dochtermann N.A. & Dingemanse N.J. 2013. Behavioral syndromes as evolutionary constraints. Behavioral Ecology, 24, 806-811.

Mueller J.C., Korsten P., Hermannstädter C., Feulner T., Dingemanse N.J., Matthysen E., van Oers K., Overveld T., Patrick S.C., Quinn J.L., Riemenschneider M., Tinbergen J.M. & Kempenaers B. 2013. Haplotype structure, adaptive history and associations with exploratory behaviour of the DRD4 region in four great tit (Parus major) populations. Molecular Ecology, In Press.

Mutzel A., Blom M.P.K., Spagopouloua F., Wright J., Dingemanse N.J. & Kempenaers B. 2013. Temporal trade-offs between nestling provisioning and defence against nest predators in blue tits. Animal Behaviour, 85, 1459-1469.

Nicolaus M., Brommer J.E., Ubels R., Tinbergen J.M. & Dingemanse N.J. 2013. Exploring patterns of variation in clutch size-density reaction norms in a wild passerine bird. Journal of Evolutionary Biology, 26, 2031-2043.

Niemelä P., Dingemanse N.J., Alioravainen N., Vainikka A. & Kortet, R. 2013. Personality pace-of-life hypothesis: testing genetic associations among personality and life-history. Behavioral Ecology, 24, 935-941.

Stuber E.F., Araya-Ajoy J.G., Mathot K.J., Nicolaus M., Wijmenga J.J., Mueller J.C. & Dingemanse N.J. 2013. Slow explorers take less risk: a problem of sampling bias in ecological studies. Behavioral Ecology, 24, 1092-1098.

Wilson A.D.M., Krause S., Dingemanse N.J. & Krause J. 2013. Network position: a key component in the characterization of social personality types. Behavioral Ecology and Sociobiology, 67, 163-173.



Carrete M., Edelaar P., Blas J., Serrano D., Potti J., Dingemanse N.J. & Tella J.L. 2012. Don't neglect pre-establishment individual selection in deliberate introductions. Trends in Ecology and Evolution, 27, 67-68.

Dall S.R.X., Gosling S.D., Brown G.D., Dingemanse N.J., Erev I., Kocher M., Schultz L., Todd P.M., Weissing F.J. & Wolf M. 2012. Variation in Decision Making. In: Evolution and the Mechanisms of Decision Making (Ed. by Hammerstein, P. & Stevens, J.R.) Cambridge: MA: MIT Press.

Dingemanse N.J., Barber I., Wright J. & Brommer J.E. 2012. Quantitative genetics of behavioural reaction norms: genetic correlations between personality and behavioural plasticity vary across stickleback populations. Journal of Evolutionary Biology, 25, 485-496.

Dingemanse N.J., Bouwman K.M., van de Pol M., van Overveld T., Patrick S.C., Matthysen E. & Quinn J.L. 2012. Variation in personality and behavioural plasticity across four populations of the great tit Parus major. Journal of Animal Ecology, 81, 116-126.

Dingemanse N.J., Dochtermann N.A. & Nakagawa S. 2012. Defining behavioural syndromes and the role of "syndrome deviation" to study its evolution. Behavioral Ecology and Sociobiology, 66, 1543-1548.

Dingemanse N.J. & Wolf M. 2012. Biological Analogs of Personality. In: Evolving the Mechanisms of Decision Making: Toward a Darwinian Decision Theory (Ed. by Hammerstein, P. & Stevens, J.R.) Cambridge, MA: MIT Press.

Mathot KJ., Wright J., Kempenaers B. & Dingemanse N.J. 2012. Adaptive strategies for managing uncertainty may explain personality-related differences in behavioural plasticity. Oikos, 121, 1009-1020.

Nicolaus M., Tinbergen J.M., Bouwman K.M., Michler S.P.M., Ubels R., Both C., Kempenaers B. & Dingemanse N.J. 2012. Experimental evidence for adaptive personalities in a wild passerine bird. Proceedings of the Royal Society of London Series B, 279, 4885-4892.

Réale D. & Dingemanse N.J. 2012. Animal personality. Encyclopedia of Life Sciences.

Sprenger D.S., Dingemanse N.J., Dochtermann N.A., Theobald J. & Walker S.P.W. 2012. Aggressive females become aggressive males in a sex-changing reef fish. Ecology Letters, 15, 986-992.



Bell, A.M., Dingemanse, N.J., Hankison, S., Langenhof, M. & Rollins, K. 2011. Early exposure to nonlethal predation risk by size-selective predators increase somatic growth and decreases size at adulthood in threespined stickleback. Journal of Evolutionary Biology, 24, 943-953.

Mathot, K., van den Hout, P., Piersma, T., Kempenaers, B., Réale, D. & Dingemanse, N.J. 2011. Disentangling the roles of frequency- versus state-dependence in generating individual differences in behavioural plasticity. Ecology Letters, 14, 1254-1262.

Mutzel, A., Kempenaers, B., Laucht, S., Dingemanse, N.J. & Dale, J. 2011. Circulating testosterone levels do not affect exploration in house sparrows: observational and experimental tests. Animal Behaviour, 81, 731-739.



Barber, I. & Dingemanse, N.J. 2010. Parasitism and the evolutionary ecology of animal personality. Philosophical Transactions of the Royal Society of London Series B, 365, 4077-4088.

Dingemanse, N.J., Dochtermann, N.A. & Wright, J. 2010. A method for exploring the structure of behavioural syndromes to allow formal comparison within and between datasets. Animal Behaviour, 79, 439-450.

Dingemanse, N.J., Edelaar, P. & Kempenaers, B. 2010. Why is there variation in glucocorticoids? Trends in Ecology and Evolution, 25, 261-262.

Dingemanse, N.J., Kazem, A.J.N., Réale, D. & Wright, J. 2010. Behavioural reaction norms: where animal personality meets individual plasticity. Trends in Ecology and Evolution, 25, 81-89.

Dingemanse, N.J. & Wolf, M. 2010. A review of recent models for adaptive personality differences. Philosophical Transactions of the Royal Society of London Series B, 365, 3947-3958.

Korsten, P., Mueller, J.C., Hermannstädter, C., Bouwman, K.M., Dingemanse, N.J., Drent, P.J., Liedvogel, M., Matthysen, E., van Oers, K., van Overveld, T., Patrick, S.C., Quinn, J.L., Sheldon, B.C., Tinbergen, J.M. & Kempenaers, B. 2010. Association between DRD4 gene polymorphism and personality in great tits: a test across four wild populations. Molecular Ecology, 19, 832-843.

Réale, D. & Dingemanse, N.J. 2010. Personality and individual social specialisation. In: Social Behaviour: Genes, Ecology and Evolution (Ed. by Szekely, T., Moore, A. & Komdeur, J.) Cambridge: Cambridge University Press.

Réale, D. & Dingemanse, N.J. 2010. Selection and evolutionary explanations for the maintenance of personality variation. In: The Evolution of Personality and Individual Differences (Ed. by Buss, D. & Hawley, P.), pp. 400-424. Oxford: Oxford University Press.

Réale, D., Dingemanse, N.J., Kazem, A.J.N. & Wright, J. 2010. Evolutionary and ecological approaches to the study of personality. Philosophical Transactions of the Royal Society of London Series B, 365, 3937-3946.

Rutten, A.L., Oosterbeek, K., Verhulst, S., Dingemanse, N.J. & Ens, B.J. 2010. Experimental evidence for interference competition in oystercatchers, Haematopus ostralegus. II. Free-living birds. Behavioral Ecology, 21, 1261-1270.



Biro, P.A. & Dingemanse, N.J. 2009. Sampling bias resulting from animal personality. Trends in Ecology and Evolution, 24, 66-67.

Both, C., Dingemanse, N.J., Drent, P.J. & Tinbergen, J.M. 2005. Pairs of extreme avian personalities have highest reproductive success. Journal of Animal Ecology, 74, 667-674.

Dijkstra, K.-D. & Dingemanse, N.J. 2000. New records of Crocothemis sanguinolenta (Burmeister, 1839) from Israel, with a cricital note on the subspecies arabica Schneider, 1982. International Journal of Odonatology, 3, 169-171.

Dingemanse,N.J. 2003. Natural Selection and Avian Personality in a Fluctuating Environment. PhD Dissertation, University of Utrecht.

Dingemanse, N.J. 2007. An evolutionary ecologist's view of how to study the persistence of genetic variation in personality. European Journal of Personality, 21, 593-596.

Dingemanse, N.J., Both, C., Drent, P.J. & Tinbergen, J.M. 2004. Fitness consequences of avian personalities in a fluctuating environment. Proceedings of the Royal Society of London Series B, 271, 847-852.

Dingemanse, N.J., Both, C., Drent, P.J., van Oers, K. & van Noordwijk, A.J. 2002. Repeatability and heritability of exploratory behaviour in great tits from the wild. Animal Behaviour, 64, 929-937.

Dingemanse, N.J., Both, C., van Noordwijk, A.J., Rutten, A.L. & Drent, P.J. 2003. Natal dispersal and personalities in great tits (Parus major). Proceedings of the Royal Society of London Series B, 270, 741-747.

Dingemanse, N.J. & de Goede, P. 2004. The relation between dominance and exploratory behavior is context-dependent in wild great tits. Behavioral Ecology, 15, 1023-1030.

Dingemanse, N.J. & Kalkman, V.J. 2008. Changing temperature regimes have advanced the phenology of Odonata in the Netherlands. Ecological Entomology, 33, 394-402.

Dingemanse, N.J., Oosterhof, C., van der Plas, F. & Barber, I. 2009. Parasitism, head morphology and the strength of morphometric correlations in sticklebacks. Biological Journal of the Linnean Society, 96, 759-768.

Dingemanse, N.J. & Réale, D. 2005. Natural selection and animal personality. Behaviour, 142, 1165-1190.

Dingemanse, N.J., van der Plas, F., Wright, J., Réale, D., Schrama, M., Roff, D.A., van der Zee, E. & Barber, I. 2009. Individual experience and evolutionary history of predation affect expression of heritable variation in fish personality and morphology. Proceedings of the Royal Society of London Series B, 276, 1285-1293.

Dingemanse, N.J., Wright, J., Kazem, A.J.N., Thomas, D.K., Hickling, R. & Dawnay, N. 2007. Behavioural syndromes differ predictably between 12 populations of stickleback. Journal of Animal Ecology, 76, 1128-1138.

Drent, P.J., Dingemanse, N.J. & van Oers, C.H.J. 2002. Functional significance and heritability in coping strategies in a wild territorial passerine: the great tit. In: Bird Research and Breeding (Ed. by Mettke-Hofmann, C. & Gansloßer, U.), pp. 25-46. Fürth: Filander Verlag.

Heg, D., Dingemanse, N.J., Lessells, C.M. & Mateman, A.C. 2000. Parental correlates of offspring sex ratio in Eurasian Oystercatchers. Auk, 117, 980-986.

Lessells, C.M., Dingemanse, N.J. & Both, C. 2002. Egg weights, egg component weights and laying gapps in great tits in relation to ambient temperature. Auk, 119, 1091-1103.

Manger, R. & Dingemanse, N.J. 2009. Adult survival of Sympecma paedisca (Brauer) during hibernation (Zygoptera: Lestidae). Odonatologica, 38, 55-59.

Nicolaus, M., Bouwman, K.M. & Dingemanse, N.J. 2008. Effect of PIT tags on the survival and recruitment of Great Tits Parus major. Ardea, 96, 286-292.

Réale, D., Reader, S.M., Sol, D., McDougall, P. & Dingemanse, N.J. 2007. Integrating temperament in ecology and evolutionary biology. Biological Reviews, 82, 291-318.

Rodenburg, J., Bastiaans, L., Schapendonk, A.H.C.M., van der Putten, P.E.L., van Ast, A., Dingemanse, N.J. & Haussmann, B.I.G. 2008. CO2-assimilation and chlorophyll fluorescence as indirect selection criteria for host tolerance against Striga. Euphytica, 160, 75-87.

Rushbrook, B.J., Dingemanse, N.J. & Barber, I. 2008. Repeatability in nest construction by male three-spined sticklebacks. Animal Behaviour, 75, 547-553.

te Marvelde, L., Meininger, P.L., Flamant, R. & Dingemanse, N.J. 2009. Age-specific density-dependent survival in Mediterranean Gulls Larus melanocephalus. Ardea, 97, 305-312.

Tieleman, B.I., Versteegh, M.A., Fries, A., Helm, B., Dingemanse, N.J., Gibbs, H.L. & Williams, J.B. 2009. Genetic modulation of energy metabolism in birds through mitochondrial function. Proceedings of the Royal Society of London Series B, 276, 1685-1693.

Tieleman, B.I., Versteegh, M.A., Helm, B. & Dingemanse, N.J. 2009. Quantitative genetics parameters show partial independent evolutionary potential for body mass and metabolism in stonechats from different populations. Journal of Zoology, 279, 129-136.

van Oers, K., Drent, P.J., Dingemanse, N.J. & Kempenaers, B. 2008. Personality is associated with extrapair paternity in great tits, Parus major. Animal Behaviour, 76, 555-563.

Versteegh, M.A., Heim, B., Dingemanse, N.J. & Tieleman, B.I. 2008. Repeatability and individual correlates of basal metabolic rate and total evaporative water loss in birds: A case study in European stonechats. Comparative Biochemistry and Physiology A-Molecular & Integrative Physiology, 150, 452-457.

Wright, J. & Dingemanse, N.J. 1999. Parents and helpers compensate for experimental changes in the provisioning effort of others in the Arabian babbler. Animal Behaviour, 58, 345-350.